We use tools from evolutionary biology, systems biology, computer science and cognitive sciences to understand how simple rules give rise to complex systems and behaviors.

Our main areas of interest are:

• Evolution of cooperation
• The role of epistasis in adaptation
• Evolution of intelligence and cognition, including learning and memory
• Effects of phenotypic plasticity on evolution
• Adaptive radiation and speciation
• Evolution of drug resistance in HIV-1
• Evolution of behavior in animals (e.g., foraging, mate selection, swarming)
• Evolution of networks (protein, gene regulatory, neural, metabolic, social)